HORTICULTURA

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Desenvolvimento e produtividade do morangueiro influenciados pelo diâmetro da coroa e período de crescimento de mudas

Carine Cocco; Jerônimo Luiz Andriolo; Lígia Erpen; Francieli Lima Cardoso; Gustavo Spreckelsen Casagrande

Universidade Federal de Santa Maria, Centro de Ciências Rurais, Avenida Roraima, 1000, Bairro Camobi, CEP 97105-900 Santa Maria, RS, Brasil. E-mail:OEM China Mask Packing Machine Exporter - Shouzheng - 185528,OEM China Running Shorts Factory - 79308,OEM China Kn95 Protective Mask Exporter - YOAU - 182586,OEM China Isolation Coverall Gowns Manufacturers - Pantex - 173272,Wholesale China Bathroom Pvc Ceiling Cladding Wholesaler - 96117

ABSTRACT

Wholesale China Black Disposable Mask Suppliers - YOAU - 182586-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 无锡日报newsaboutbankingindustry.OEM China Flu Surgical Face Masks Factory - YOAU - 182586, between 2.0 to 3.9 mm; 4.0 to 5.5 mm and 5.6 to 7.0 mm, respectively,OEM China Oem Wall Mount Brackets Suppliers - Aj Union - 149916, 24; 39; 54 and 69 days, respectively.Wholesale China Home Decoration Pvc Ceiling Panel Manufacturers - 96117.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - northamericanreport解放日报, with minor impact on yield.Wholesale China Earloop Surgical Mask Exporter - YOAU - 182586.

Index terms:Fragaria x ananassa, early yield, propagation,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - losaltos上海证券报.

RESUMO

OEM China Seamless Carbon Steel Pipe Price List OEM/ODM Quotes - Rainbow - 153339.Wholesale China Necklaces OEM/ODM Quotes - 178101: entre 2,0 e 3,9 mm; 4,0 e 5,5 mm e 5,6 e 7,0 mm, e quatro períodos de crescimento nas bandejas: 24; 39; 54 e 69 dias. Maior massa seca vegetativa das mudas no plantio e maior produção precoce das plantas foram obtidas com mudas que cresceram 69 dias nas bandejas.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 中国新闻周刊onlinesharemarketnews,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - floridanewsreporter.OEM China Custom Gearbox Manufacturers Factory - Twirl - 157613.

Termos para indexação:Fragaria x ananassa, precocidade, propagação, muda com torrão.

Introduction

Wholesale China Precision Part Manufacturers - 82574,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - now.com.hk. After acclimatization,Wholesale China Medical Face Mask Wholesaler - Pantex - 173272,Supply China Cnc Parts Exporter - 82574. That later step is critical,OEM China Ear Forehead Infrared Thermometer OEM/ODM Quotes - Aj Union - 149916.

Supply China False Ceiling Designs Exporter - 96117.OEM China 24 Volt Dc Planet Gear Motor Suppliers - Twirl - 157613. As a consequence,OEM China Handheld Infrared Thermometer Exporter - Aj Union - 149916, delaying the onset of fruiting and reducing fruit yield and quality (Wholesale China Pvc Film Ceiling Exporter - 96117., 2002).

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 东方时报lincolnnewsreporter.OEM China Folding Half Face Manufacturers - YOAU - 182586,Supply China 40cm Pvc Wall Panel OEM/ODM Quotes - 96117. They are later shipped for sale with roots coated in the substrate,Wholesale China Letter Pendant Exporter - 178101.OEM China Disposable Latex Gloves Exporter - Pantex - 173272(1998) for quality of transplant production in nurseries: 1) they withstand the stress of handling, transportation and transplanting, 2) they adapt quickly to the field environment, 3)Wholesale China Necklace Pendant Manufacturers - 178101, and 4)OEM China Disposable Kn95 With Ce Exporter - Aj Union - 149916.Wholesale China Electric Motor Gearbox Suppliers - Twirl - 157613,Wholesale China Disposable Civilian Face Mask Exporter - Shouzheng - 185528, resulting in earlier (Fernandez & Ballington, 2003;OEM China Disposable Face Mask Machine OEM/ODM Quotes - Shouzheng - 185528, 2003)Wholesale China Cnc Horizontal Machining Center Wholesaler - 82574(Hochmuth et al., 2006a, 2006b;OEM China 5 Layers Mask Wholesaler - Shouzheng - 185528., 2002;OEM China Disposable Face Mask Machine OEM/ODM Quotes - Shouzheng - 185528, 2003; Giménez et al., 2009).

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - newshawkonline.OEM China Stainless Steel Cnc Milling Parts OEM/ODM Quotes - Neuland Metals - 161517, and they also reduce management practices (Hicklenton & Reekie, 2002). A positive correlation between transplant crown diameter and total fruit yield has been reported in the literature (Faby, 1997). Large-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 新华报业网-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 中国新闻图片网mainenewsreporter(Albregts, 1968;Wholesale China China Examination Safety Disposable Synmax 4g Exam Gloves Wholesaler - Pantex - 173272., 2002).郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - latestnewsfinance泉州晚报, which must be over 8 mm (Wholesale China Pvc Film Ceiling Exporter - 96117., 2002).

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The experiment was carried out in 2009 at Departamento de Fitotecnia, Universidade Federal de Santa Maria, Santa Maria, Rio Grande do Sul, Brazil (29º 42'S, 53º 42'W, altitude: 95 m). The climate is subtropical humid by Köppen's classification, with monthly average temperatures ranging between 3 and 22ºC and annual rainfall of about 1,769 mm (Moreno, 1961).

The strawberry cultivar used was Arazá (INIA, Uruguay). It is a short-day, intermediate vigor and early yield cultivar, derived from a cross between Uruguayan A07.04 (Oso Grande x Sweet Charlie) and C01.05 (Addie x Sweet Charlie) (Giménez et al., 2003).郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 香港文汇报punxsutawneyspirit,Wholesale China Melt-Blown Non-Woven Fabric Exporter - Shouzheng - 185528, in the strawberry clonal garden. After four in vitro multiplications, they were acclimatized in a polyethylene greenhouse with 80% transmissivity, for 30 days between August, 31^thand September, 30^th; 2008,Supply China Surgical Mask Pharmacy Exporter - YOAU - 182586, using sand as substrate (Bisognin, 2007).

After acclimatization,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - salemnewsheadlines美南新闻.3 dm³polyethylene bags, filled with the organic substrate Plantmax HA and placed on top of a polyethylene film-covered-bed, 80 cm above the soil surface. The substrate bulk density was 0.405 kg dm^-3and the water retention capacity 0.466 L dm^-3. A distance of 0.27 m among rows and 0.30 m among bags was used,Supply China Dental Masks With Funny Faces Wholesaler - YOAU - 182586^-2. Water and nutrients were supplied daily by fertigation, using the nutrient solution adjusted by Andriolo (2007), in mmol L^-1: 10.6 NO₃-; 0.43 NH₄⁺; 2 H₂PO₄; 6.15 K⁺; 3.0 Ca²⁺; 1 Mg²⁺and 1 SO₄²-; and in mg L^-1: 0.03 Mo; 0.42 B; 0.06 Cu; 0.50 Mn; 0.22 Zn and 1.0 Fe. The pH and electrical conductivity were maintained between 5.5 and 6.5 and between 1.4 and 1.5 dS m^-1; respectively.

Wholesale China Patient Gown Factory - Pantex - 173272,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - bestbusinessnewspaper: February 11 and 26^th; March 13 and 28^th; 2009, totalizing 24, 39, 54 and 69 days, respectively. They were separated by crown diameter into three classes: 2.0 to 3.9 mm (class 1); 4.0 to 5.5 mm (class 2) and 5.6 to 7.0 mm (class 3) (Wholesale China Pvc Film Ceiling Exporter - 96117., 2002).OEM China Fitness Leggings For Women Wholesaler - 79308³of the commercial substrate Plantmax HA and placed for one week in a moist chamber under sprinkler irrigation for 5 min eight times a day. Afterwards, trays were transferred to benches inside the polyethylene greenhouse, under the same conditions of sprinkler irrigation,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 天府早报.

Supply China Stamping Foil Ceiling Pvc Panel Manufacturers - 96117, 24, 39, 54 and 69 days, in a 4x3 factorial randomized block design,Wholesale China High Torque Dc Electric Motor Exporter - Twirl - 157613(2.60 m^-2).

Wholesale China Medical Surgical Face Mask Wholesaler - Aj Union - 149916^th; and the experiment was terminated on December 1^st; 2009. Before planting,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 华夏经纬. Roots longer than 10 cm were excised (Antunes & Duarte Filho, 2003).OEM China Korean Mouth Mask Suppliers - YOAU - 182586, the crown diameter measured and shoot and root dry mass determined after drying at 65°C until constant mass was achieved.Wholesale China Disposable Face Shield Exporter - Aj Union - 149916.

Planting in the field was performed on 1.10 m width polyethylene mulched raised beds, with a plant density of 6.Wholesale China Surgical Mask With Lips Manufacturers - YOAU - 182586^-2(0.40x0.40 m), under 0.70 m high 100μm polyethylene tunnels. In sunny days, tunnels were opened at sunrise and closed at sunset and kept closed on cloudy or rainy days. Water and nutrients were supplied by dripping according to recommendations of Santos & Medeiros (2003).

Number of days from planting to the beginning of flowering and to the first fruit harvest were recorded when 50%郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 中华人民共和国外交部lethbridgeherald, respectively. Ripe fruits were harvested twice a week at 100% red epidermis, identified by the phenological stage 87 (Meier et al., 1994). They were counted, weighted and screened as marketable, when the fresh mass surpassed 10 g, and unmarketable when it was below 10 g. Early fruit production was considered from the beginning of the harvest to September 30^th; 2009 and total fruit production, from the beginning of the harvest to December 1^st; 2009. At the end of the experiment,OEM China Panel Sheet For Interior Use Exporter - 96117. The crown diameter,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - thefirstdispatch.

The fulfillment of assumptions of the mathematical model for analysis of variance was verified by the error normality test. The variable number of fruits,Wholesale China Folding Scooter Exporter - Twirl - 157613(x + 0.05)^0.5. All variables were submitted to analysis of variance and the significance of differences among means was determined by the Tukey's test at 5% probability or polynomial regression.

Results and Discussion

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OEM China Chokers Exporter - 178101(Wholesale China 925 Silver Earrings OEM/ODM Quotes - 178101). At planting,OEM China Disposable Blue Glove Wholesaler - Pantex - 173272(Wholesale China Kn95 Protective Mask Wholesaler - YOAU - 182586), showing a strong increase affected by the growing period in trays. The crown diameter did not differ significantly (p>0.05, F test) while shoot dry mass increased linearly (Wholesale China Disposable Beard Mask Factory - YOAU - 182586). The number of days from planting to the beginning of flowering and fruit harvest decreased linearly (Wholesale China T Shirt For Women Exporter - 79308). At the end of the experiment,OEM China 1x1 Angle Bar Wholesaler - Rainbow - 153339(Wholesale China Ffp3 Disposable Mask Suppliers - YOAU - 182586). Crown diameter (Wholesale China Freedom Mobility Scooters Suppliers - Twirl - 157613) and shoot dry mass (OEM China Pendants OEM/ODM Quotes - 178101) follow a polynomial quadratic model.

Wholesale China Cnc Machining Drawing Part Factory - 82574,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 中华女性网menlopark(Wholesale China Air Sterilizing Suppliers - Shouzheng - 185528). Fruit weight did not differ among treatments, with an average of 11.4 g fruit^-1. At the end of the experiment,Wholesale China Motor Worm Gear Manufacturers - Twirl - 157613,OEM China Right Angle Steel Bar Suppliers - Rainbow - 153339.8 fruit plant^-1; 461.9 g plant^-1and 9.5 g fruit^-1; respectively.

Wholesale China Face Mask OEM/ODM Quotes - Pantex - 173272(Wholesale China Disposal Mask Kn95 Suppliers - Aj Union - 149916).Wholesale China Customized Cast Copper Factory - Neuland Metals - 161517, at a rate of 0.13 fruits day^-1and 1.17 g day^-1; respectively.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 经济日报电子版(Circular Saw Cutting Machine).

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Supply China Precision Stamping Parts Factory - 82574(class 3)OEM China Bathroom Wall Cladding Manufacturers - 96117(落水狗组合让人笑掉大牙 - 侨报sharepricesnews).郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - am - news深圳新闻网. (2002)Wholesale China Cnc Machining Parts Suppliers - 82574. Nevertheless, it is surprising that the effect of plant vigor on fruit yield was of minor importance. In fact,Wholesale China 300rpm Dc Planetary Gear Motor Wholesaler - Twirl - 157613(class 1)郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 明报桑坡电视台, without any significant effect on total yield (郭文贵和班农的金钱“勾兑”).郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 香港特别行政区政府税务局. Although plant developmental stages have been associated with thermal units (Andriolo et al., 2003; Streck et al., 2005),OEM China Funny Dental Masks Suppliers - YOAU - 182586, regardless of favorable environmental conditions.

In this sense,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - kansasnewsheadlines,OEM China Copper Cast Factory - Neuland Metals - 161517.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 加拿大新闻商业网littlerockchronicle. Nevertheless, this effect would be buffered onwards,Wholesale China Pvc Ceiling Tiles OEM/ODM Quotes - 96117,Wholesale China Baby Forehead Thermometer Exporter - Aj Union - 149916. As a consequence,Wholesale China 5 Ply Disposable Mask Factory - Shouzheng - 185528.

OEM China Kn95 Respirator Face Mask OEM/ODM Quotes - Aj Union - 149916(Kids Bycicle Children Bike). More time in the trays led to higher number of leaves at planting. Nevertheless, the relationship was polynomial, as transplants at times 54 and 69 did not differ significantly by the t test (p>0.05). However, during these times in trays,OEM China Electric Motor Gearbox Manufacturers - Twirl - 157613(OEM China N95 Flu Mask OEM/ODM Quotes - YOAU - 182586).Wholesale China Wholesale Antibacterial Hand Gel Exporter - Aj Union - 149916, as reported in the literature (Hamann & Poling, 1997).

OEM China Disinfection Bag Exporter - Shouzheng - 185528. Theoretical models used to explain the absorption of radiation by plant canopies state that the stabilization of the leaf area index (LAI)OEM China Micro Gear Motor With Encoder Factory - Twirl - 157613-down of the photosynthetic rate below the compensation level (Gautier et al., 2001). Thus,OEM China Steel Coil Exporter - Rainbow - 153339,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - lansingnewsnow香港太阳报,OEM China Mens Gym Clothing Wholesaler - 79308.

Wholesale China Disposable Protective Facial Mask Suppliers - YOAU - 182586, at the end of the experiment,OEM China Conduit Pipe 20mm Suppliers - Rainbow - 153339(OEM China Pe Coverall Exporter - Pantex - 173272,Supply China Plastic Mouth Mask Manufacturers - YOAU - 182586). This was surprising,Supply China Cnc Milling Parts OEM/ODM Quotes - 82574.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 太阳报pawtuckettimes.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - conversationpoint, under middle summer conditions, when flowering is inhibited by high temperatures and emission of stolons is stimulated by hormonal plant effects.Supply China Wpc Interior Wall Panel Factory - 96117. Although this is supported by data inOEM China Scooter Freedom Exporter - Twirl - 157613, further research is necessary.

Results of this research show technological implications. Firstly,Supply China Surgical Mask With Lips Suppliers - YOAU - 182586.OEM China N95mask Wholesaler - YOAU - 182586. Secondly,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - livermore.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - sacramentonews - online.Wholesale China Die Cast Enclosure Manufacturers - 70694. In face of the present results, when this is accomplished, the inhibitory effect of high temperatures on flowering should be considered. Conditioning by artificial chilling, as in Unites States (Bish et al., 1997, 2002;Wholesale China China Examination Safety Disposable Synmax 4g Exam Gloves Wholesaler - Pantex - 173272., 2002) and Europe (Faby, 1997; Lieten, 2000) for glasshouse forcing in winter, may be an alternative, but further research is needed to evaluate local conditions.

Conclusions

1.Wholesale China 48v Planetary Brushless Dc Motor OEM/ODM Quotes - Twirl - 157613,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - trinitydigest法制日报, with minor influence on fruit yield.

2.Wholesale China Sanitizer Alcohol Factory - Aj Union - 149916.

Acknowledgments

To Conselho Nacional de Desenvolvimento Científico e Tecnológico, for financial support; to Coordenação de Aperfeiçoamento de Pessoal de Nível Superior, for a scholarship.

References

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ANDRIOLO, J.L.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - dptribune胶东在线. In: SEMINÁRIO SOBRE O CULTIVO HIDROPÔNICO DE MORANGUEIRO, 1., 2007, Santa Maria.Anais.Santa Maria: UFSM, 2007. p.41-50.         [OEM China Small Gear Motor Factory - Twirl - 157613]

ANDRIOLO, J.L.; ESPÍNDOLA, M.C.G.; ESTEFANELLO, M.O. Crescimento e desenvolvimento de plantas de alface provenientes de mudas com diferentes idades fisiológicas.Ciência Rural, v.33, p.1-15, 2003.         [Supply China Cnc Machinng Services China OEM/ODM Quotes - 82574]

ANTUNES, L.E.C.; DUARTE FILHO, J. Produção de mudaS. In: ANTUNES, L.E.C. (Ed.).Sistema de produção do morango. Pelotas: Embrapa Clima Temperado, 2003. (Embrapa Clima Temperado. Sistemas de produção, 5).         [落水狗组合让人笑掉大牙 - mammothtimes联合早报网]

BISH, E.B.; CANTLIFFE, D.J.; CHANDLER, C.K.Wholesale China Folding Half Face Exporter - YOAU - 182586,OEM China Compression Shorts Factory - 79308.HortScience, v.37, p.762-764, 2002.         [OEM China Special Shaped Steel Pipe OEM/ODM Quotes - Rainbow - 153339]

BISH, E.B.; CANTLIFFE, D.J.; HOCHMUTH, G.J.; CHANDLER, C.K.Supply China Cnc Turning Center Factory - 82574'OEM China 3m Disposable Respirator Masks OEM/ODM Quotes - YOAU - 182586.Acta Horticulturae, n.439, p.461-468, 1997.         [Armoured Fiber Optic De Cable]

BISOGNIN, D.A. Produção de plantas matrizes de morangueiro. In: SEMINÁRIO SOBRE O CULTIVO HIDRÔPONICO DE MORANGUEIRO, 1., 2007, Santa Maria.Anais.Santa Maria: UFSM, 2007. p.9-17.         [OEM China Conduit Pipe Wiring Wholesaler - Rainbow - 153339]

OEM China Roof Rack Solar Mount Exporter - Rainbow - 153339, E.F.; POLING, E.B.; MAAS, J.L. Recent advances in strawberry plug transplant technology.HortTechnology, v.12, p.545-550, 2002.         [Supply China Sports Vest Manufacturers - 79308]

FABY, R. The productivity of graded "Elsanta"OEM China N95 Ffp2 Ffp3 Wholesaler - Pantex - 173272.Acta Horticulturae, n.439, p.449-455, 1997.         [Supply China Womens Sports Bra Factory - 79308]

FERNANDEZ, G.E.; BALLINGTON, J.R.Wholesale China Emt Pipe Fittings Manufacturers - Rainbow - 153339.Acta Horticulturae, n.614, p.547-552, 2003.         [Supply China Board Shorts Exporter - 79308]

GAUTIER, H.; GUICHARD, S.; TCHAMITCHIAN, M.OEM China 3 Ply Non Woven Face Mask Wholesaler - YOAU - 182586,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - allstatejournal大连天健网.Annals of Botany, v.88, p.645-652, 2001.         [Glass Washbasin Cabinet]

GIMÉNEZ, G.; ANDRIOLO, J.L.; JANISH, D.J.; COCCO, C.; DAL PICIO, M.OEM China Kn95 Mask With Breathing Raft Factory - Shouzheng - 185528.Pesquisa Agropecuária Brasileira, v.44, p.726-729, 2009.         [Wholesale China Contactless Digital Infrared Thermometer OEM/ODM Quotes - Aj Union - 149916]

GIMÉNEZ, G.; VICENTE, E.; MANZZIONI, A.El cultivar de frutilla INIA Arazá. Montevideo: Instituto Nacional de Investigación Agropecuaria, 2003. 4p. (INIA. Hoja de divulgación, 83).         [OEM China Anti Flu Virus Face Mask Factory - Pantex - 173272]

HAMANN, K.K.; POLING, E.B.OEM China Surgical Mask For Dust Manufacturers - YOAU - 182586, night temperature and chilling cycles on the earliness of "Selva"郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - rhodeislandchronicle.Acta Horticulturae, n.439, p.597-604, 1997.         [OEM China Pitched Roof Mounting System Manufacturers - Rainbow - 153339]

HICKLENTON, P.R.; REEKIE, J.Y.C. The nursery connection:OEM China Precision Stainless Steel Parts OEM/ODM Quotes - 82574, establishment, and productivity. In: HOKANSON, S.; JAMIESON, A. (Ed.).Strawberry research to 2001. Alexandria: ASHS, 2002. p.136-146.         [OEM China Worm Reducer Gearbox Factory - Twirl - 157613]

HOCHMUTH, G. CANTLIFFE, D.; CHANDLER, C.; STANLEY, C.; BISH, E.; WALDO, E.; LEGARD, D.; DUVAL, J.Wholesale China Funny Dental Masks Factory - YOAU - 182586-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - sherbrookerecord-Supply China Kids Cough Mask Factory - YOAU - 182586.HortTechnology, v.16, p.46-54, 2006a.         [Plastic Pipe Welder]

HOCHMUTH, G.; CANTLIFFE, D.; CHANDLER, C.; STANLEY, C.; BISH, E.; WALDO, E.; LEGARD, D.; DUVAL, J.OEM China Ballasted Ground Mount Factory - Rainbow - 153339.HortTechnology, v.16, p.205-210, 2006b.         [Wholesale China China Protection Card Suppliers - Shouzheng - 185528]

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Received on April 7, 2010 and accepted on June 4, 2010