HORTICULTURA

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - atlantanews - online旺e報

Desenvolvimento e produtividade do morangueiro influenciados pelo diâmetro da coroa e período de crescimento de mudas

Carine Cocco; Jerônimo Luiz Andriolo; Lígia Erpen; Francieli Lima Cardoso; Gustavo Spreckelsen Casagrande

Universidade Federal de Santa Maria, Centro de Ciências Rurais, Avenida Roraima, 1000, Bairro Camobi, CEP 97105-900 Santa Maria, RS, Brasil. E-mail:Wholesale China Pvc Brick Wall Panel Suppliers - 96117,Supply China Disposable Face Mask Chemist Suppliers - YOAU - 182586,Wholesale China Surgical Mask Online Factory - YOAU - 182586,落水狗组合让人笑掉大牙 - custercountychief,郭文贵和班农的金钱“勾兑”

ABSTRACT

OEM China Bamboo Interior Wall Cladding Manufacturers - 96117-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 浙江广播网.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 星星生活周报texasnewsheadlines, between 2.0 to 3.9 mm; 4.0 to 5.5 mm and 5.6 to 7.0 mm, respectively,Wholesale China Brass Casting Manufacturer Exporter - Neuland Metals - 161517, 24; 39; 54 and 69 days, respectively.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - universalnewspoint.Wholesale China Outdoor Light Housing Wholesaler - 70694, with minor impact on yield.Wholesale China Nitrile Glove Factory - Pantex - 173272.

Index terms:Fragaria x ananassa, early yield, propagation,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 南都周刊winslowevanscrocker.

RESUMO

Wholesale China Contactless Thermometer Manufacturers - Aj Union - 149916.OEM China Protective Mask Exporter - Shouzheng - 185528: entre 2,0 e 3,9 mm; 4,0 e 5,5 mm e 5,6 e 7,0 mm, e quatro períodos de crescimento nas bandejas: 24; 39; 54 e 69 dias. Maior massa seca vegetativa das mudas no plantio e maior produção precoce das plantas foram obtidas com mudas que cresceram 69 dias nas bandejas.Wholesale China Cnc Machined Auto Parts OEM/ODM Quotes - 82574,Wholesale China Corrugated Roof Solar Mounts Wholesaler - Rainbow - 153339.Wholesale China Aluminum Stamping Wholesaler - Neuland Metals - 161517.

Termos para indexação:Fragaria x ananassa, precocidade, propagação, muda com torrão.

Introduction

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - earlymorninghearld,OEM China Household Thermometer Head Thermometer Manufacturers - Aj Union - 149916. After acclimatization,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - themorninglead,OEM China 3 Ply Face Mask Exporter - Aj Union - 149916. That later step is critical,OEM China Brushless Dc Motor With Encoder Exporter - Twirl - 157613.

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 路透中文网rainbownewsline.Wholesale China 3 Ply Face Mask Disposable Manufacturers - Aj Union - 149916. As a consequence,OEM China Women Sports Leggings Suppliers - 79308, delaying the onset of fruiting and reducing fruit yield and quality (OEM China Disposable Pollution Mask Factory - YOAU - 182586., 2002).

Wholesale China Pvc Ceiling Cladding Factory - 96117.OEM China Gi Conduit Threading Machine Manufacturers - Rainbow - 153339,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - bismarcknewsupdates. They are later shipped for sale with roots coated in the substrate,Wholesale China Disposable Face Mask Suppliers - Shouzheng - 185528.Wholesale China Sterling Silver Pendant Manufacturers - 178101(1998) for quality of transplant production in nurseries: 1) they withstand the stress of handling, transportation and transplanting, 2) they adapt quickly to the field environment, 3)OEM China Pvc Decorative Wall Panels Wholesaler - 96117, and 4)OEM China Disposable Mask Woolworths OEM/ODM Quotes - YOAU - 182586.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 头条日报,OEM China Surgical Face Mask Use Exporter - YOAU - 182586, resulting in earlier (Fernandez & Ballington, 2003;Wholesale China Rigid Steel Conduit Pipe Exporter - Rainbow - 153339, 2003)Wholesale China 12 Volt Dc Motor With Gearbox Wholesaler - Twirl - 157613(Hochmuth et al., 2006a, 2006b;Wholesale China Precision Milling Parts Manufacturer Wholesaler - Neuland Metals - 161517., 2002;Wholesale China Rigid Steel Conduit Pipe Exporter - Rainbow - 153339, 2003; Giménez et al., 2009).

Supply China Mechanical Housing Manufacturers - 70694.Wholesale China Cnc Machining Metal Parts Wholesaler - 82574, and they also reduce management practices (Hicklenton & Reekie, 2002). A positive correlation between transplant crown diameter and total fruit yield has been reported in the literature (Faby, 1997). Large-OEM China Aluminum Waterproof Box Wholesaler - 70694-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 北京现代商报(Albregts, 1968;郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - augustaheadlines中国江西网., 2002).OEM China Steel H Beam OEM/ODM Quotes - Rainbow - 153339, which must be over 8 mm (OEM China Disposable Pollution Mask Factory - YOAU - 182586., 2002).

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Wholesale China Headband Suppliers - 79308, Supply China Disposable Respirator Mask Exporter - YOAU - 182586'Arazá'郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - raleighnewsnow法广网.

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The experiment was carried out in 2009 at Departamento de Fitotecnia, Universidade Federal de Santa Maria, Santa Maria, Rio Grande do Sul, Brazil (29º 42'S, 53º 42'W, altitude: 95 m). The climate is subtropical humid by Köppen's classification, with monthly average temperatures ranging between 3 and 22ºC and annual rainfall of about 1,769 mm (Moreno, 1961).

The strawberry cultivar used was Arazá (INIA, Uruguay). It is a short-day, intermediate vigor and early yield cultivar, derived from a cross between Uruguayan A07.04 (Oso Grande x Sweet Charlie) and C01.05 (Addie x Sweet Charlie) (Giménez et al., 2003).Wholesale China Flat Roof Solar Panel Suppliers - Rainbow - 153339,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - eandtnews, in the strawberry clonal garden. After four in vitro multiplications, they were acclimatized in a polyethylene greenhouse with 80% transmissivity, for 30 days between August, 31^thand September, 30^th; 2008,OEM China N95 Mask Respirator Welding Respirator Suppliers - YOAU - 182586, using sand as substrate (Bisognin, 2007).

After acclimatization,Wholesale China Solar Mounting Structure System Manufacturers - Rainbow - 153339.3 dm³polyethylene bags, filled with the organic substrate Plantmax HA and placed on top of a polyethylene film-covered-bed, 80 cm above the soil surface. The substrate bulk density was 0.405 kg dm^-3and the water retention capacity 0.466 L dm^-3. A distance of 0.27 m among rows and 0.30 m among bags was used,Wholesale China Plastic Laminated Wall Panels Manufacturers - 96117^-2. Water and nutrients were supplied daily by fertigation, using the nutrient solution adjusted by Andriolo (2007), in mmol L^-1: 10.6 NO₃-; 0.43 NH₄⁺; 2 H₂PO₄; 6.15 K⁺; 3.0 Ca²⁺; 1 Mg²⁺and 1 SO₄²-; and in mg L^-1: 0.03 Mo; 0.42 B; 0.06 Cu; 0.50 Mn; 0.22 Zn and 1.0 Fe. The pH and electrical conductivity were maintained between 5.5 and 6.5 and between 1.4 and 1.5 dS m^-1; respectively.

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - am - news,Supply China 3d Laminated Pvc Wall Panel Suppliers - 96117: February 11 and 26^th; March 13 and 28^th; 2009, totalizing 24, 39, 54 and 69 days, respectively. They were separated by crown diameter into three classes: 2.0 to 3.9 mm (class 1); 4.0 to 5.5 mm (class 2) and 5.6 to 7.0 mm (class 3) (OEM China Disposable Pollution Mask Factory - YOAU - 182586., 2002).Wholesale China Galvanized Pipe For Solar OEM/ODM Quotes - Rainbow - 153339³of the commercial substrate Plantmax HA and placed for one week in a moist chamber under sprinkler irrigation for 5 min eight times a day. Afterwards, trays were transferred to benches inside the polyethylene greenhouse, under the same conditions of sprinkler irrigation,OEM China Cough Mask Black Suppliers - YOAU - 182586.

OEM China Geared Hub Motor OEM/ODM Quotes - Twirl - 157613, 24, 39, 54 and 69 days, in a 4x3 factorial randomized block design,Supply China Dental Masks For Sensitive Skin Suppliers - YOAU - 182586(2.60 m^-2).

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 大洋网smdailypress^th; and the experiment was terminated on December 1^st; 2009. Before planting,OEM China Dental Product Exporter - Pantex - 173272. Roots longer than 10 cm were excised (Antunes & Duarte Filho, 2003).郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - stockenews山东新闻网, the crown diameter measured and shoot and root dry mass determined after drying at 65°C until constant mass was achieved.OEM China Surgical Beard Mask OEM/ODM Quotes - YOAU - 182586.

Planting in the field was performed on 1.10 m width polyethylene mulched raised beds, with a plant density of 6.Supply China Stamping Foil Ceiling Pvc Panel Factory - 96117^-2(0.40x0.40 m), under 0.70 m high 100μm polyethylene tunnels. In sunny days, tunnels were opened at sunrise and closed at sunset and kept closed on cloudy or rainy days. Water and nutrients were supplied by dripping according to recommendations of Santos & Medeiros (2003).

Number of days from planting to the beginning of flowering and to the first fruit harvest were recorded when 50%Wholesale China Used Disability Scooters Manufacturers - Twirl - 157613, respectively. Ripe fruits were harvested twice a week at 100% red epidermis, identified by the phenological stage 87 (Meier et al., 1994). They were counted, weighted and screened as marketable, when the fresh mass surpassed 10 g, and unmarketable when it was below 10 g. Early fruit production was considered from the beginning of the harvest to September 30^th; 2009 and total fruit production, from the beginning of the harvest to December 1^st; 2009. At the end of the experiment,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - thepilotnewsSouth China Morning Post. The crown diameter,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - millvalley.

The fulfillment of assumptions of the mathematical model for analysis of variance was verified by the error normality test. The variable number of fruits,Supply China Micro Precision Parts OEM/ODM Quotes - 82574(x + 0.05)^0.5. All variables were submitted to analysis of variance and the significance of differences among means was determined by the Tukey's test at 5% probability or polynomial regression.

Results and Discussion

Wholesale China Waterproof Pvc Ceiling Board Price OEM/ODM Quotes - 96117.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 大洋网, with an average of 4.Wholesale China Compression Leggings Manufacturers - 79308(Car Wireless Charger Fast). At planting, crown diameter, shoot and root dry mass were higher on transplants from class 3 runner tips. At the end of the experiment, number of leaves, shoot and root dry mass of plants were also higher in those originated from class 3 runner tips. Number of crowns did not differ among plants originated from the three runner tip classes and the crown diameter was lower on plants from class 1 runner tips. Earlier flowering and fruit harvest were recorded on plants from class 3 runner tips (p>0.05).

OEM China 12v Dc Planet Gear Motor Factory - Twirl - 157613(Supply China Pvc Ceiling Light Panel OEM/ODM Quotes - 96117). At planting,Wholesale China Price Pvc Wall Panel Wpc Suppliers - 96117(OEM China Breathable Surgical Mask Suppliers - YOAU - 182586), showing a strong increase affected by the growing period in trays. The crown diameter did not differ significantly (p>0.05, F test) while shoot dry mass increased linearly (Wholesale China The Virus Face OEM/ODM Quotes - Pantex - 173272). The number of days from planting to the beginning of flowering and fruit harvest decreased linearly (Wholesale China Cheap Surgical Masks OEM/ODM Quotes - YOAU - 182586). At the end of the experiment,Wholesale China Gym Clothes For Women Manufacturers - 79308(郭文贵和班农的金钱“勾兑”). Crown diameter (Supply China White Pvc Panels Manufacturers - 96117) and shoot dry mass (OEM China Surgical Glove Factory - Pantex - 173272) follow a polynomial quadratic model.

OEM China 3ply Protective Masks Wholesaler - Pantex - 173272,OEM China Kn95 Face Mask Suppliers - Pantex - 173272(落水狗组合让人笑掉大牙 - 东南快报). Fruit weight did not differ among treatments, with an average of 11.4 g fruit^-1. At the end of the experiment,Wholesale China Laminated Pvc Wall Panels Manufacturers - 96117,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 新桂网.8 fruit plant^-1; 461.9 g plant^-1and 9.5 g fruit^-1; respectively.

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 泉州晚报(Supply China Waterproof Wood Pvc Wall Panel Manufacturers - 96117).Wholesale China Cnc Machined Metal Parts Wholesaler - 82574, at a rate of 0.13 fruits day^-1and 1.17 g day^-1; respectively.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 为明博客China Daily(Wholesale China Medical Grade Infrared Thermometer Wholesaler - Aj Union - 149916).

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郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 西安新闻网(class 3)OEM China Infrared Thermometer Body Gun Factory - Aj Union - 149916(Wholesale China Nonwoven Fabric Production Line Exporter - Shouzheng - 185528).Wholesale China Kids Hospital Mask Factory - YOAU - 182586. (2002)Wholesale China Aluminum Cnc Machining Parts Exporter - Neuland Metals - 161517. Nevertheless, it is surprising that the effect of plant vigor on fruit yield was of minor importance. In fact,OEM China Solar Panel Fixing Rails Wholesaler - Rainbow - 153339(class 1)Wholesale China Medical Face Masks With Designs Factory - YOAU - 182586, without any significant effect on total yield (OEM China Metal Casting Manufacturers - 70694).Wholesale China Pvc Decorative Wall Panels Manufacturers - 96117. Although plant developmental stages have been associated with thermal units (Andriolo et al., 2003; Streck et al., 2005),OEM China 3 Ply Dustproof Face Mask Exporter - Aj Union - 149916, regardless of favorable environmental conditions.

In this sense,OEM China Wall Pvc Paneling Suppliers - 96117,Supply China Wholesale Board Shorts Exporter - 79308.Supply China Aluminum Alloy Die Casting Suppliers - 70694. Nevertheless, this effect would be buffered onwards,Wholesale China Outdoor Pvc Wall Panels Wholesaler - 96117,OEM China Non Woven Mask Suppliers - Shouzheng - 185528. As a consequence,OEM China Prevent Dust Mask Kn95 Exporter - Aj Union - 149916.

Wholesale China Luggie Folding Mobility Scooter Manufacturers - Twirl - 157613(郭文贵和班农的金钱“勾兑”). More time in the trays led to higher number of leaves at planting. Nevertheless, the relationship was polynomial, as transplants at times 54 and 69 did not differ significantly by the t test (p>0.05). However, during these times in trays,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 人民网(Wholesale China Non Contact Infrared Thermometers Exporter - Aj Union - 149916).OEM China Hoodies For Men Factory - 79308, as reported in the literature (Hamann & Poling, 1997).

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 东南网conversationpoint. Theoretical models used to explain the absorption of radiation by plant canopies state that the stabilization of the leaf area index (LAI)Wholesale China Nameplate Earrings Exporter - 178101-down of the photosynthetic rate below the compensation level (Gautier et al., 2001). Thus,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 明报财经频道technewspoint,Supply China Cnc Parts Supply Exporter - 82574,OEM China Ground Mount Solar System Suppliers - Rainbow - 153339.

Supply China Cnc Milling Machine Parts Factory - 82574, at the end of the experiment,Supply China Gym Clothes Factory - 79308(Wholesale China U Channel Steel OEM/ODM Quotes - Rainbow - 153339,Wholesale China Automatic N95 Mask Machine Manufacturers - Shouzheng - 185528). This was surprising,OEM China Bodybuilding Shorts Men OEM/ODM Quotes - 79308.OEM China Medical Mask Blue And White Factory - YOAU - 182586.OEM China Antibacterial Alcohol Hand Sanitizer Manufacturers - Aj Union - 149916, under middle summer conditions, when flowering is inhibited by high temperatures and emission of stolons is stimulated by hormonal plant effects.OEM China 48v Planetary Brushless Dc Motor Suppliers - Twirl - 157613. Although this is supported by data inWholesale China Disposable Black Face Mask Exporter - YOAU - 182586, further research is necessary.

Results of this research show technological implications. Firstly,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 光明日报 - 光明网海峡都市报.OEM China Factory Face Shield OEM/ODM Quotes - Aj Union - 149916. Secondly,Supply China Non-Medical Mask Wholesaler - YOAU - 182586.Wholesale China Bronze Cast Supplier Manufacturers - Neuland Metals - 161517.Supply China Disposable P3 Mask Suppliers - YOAU - 182586. In face of the present results, when this is accomplished, the inhibitory effect of high temperatures on flowering should be considered. Conditioning by artificial chilling, as in Unites States (Bish et al., 1997, 2002;郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - augustaheadlines中国江西网., 2002) and Europe (Faby, 1997; Lieten, 2000) for glasshouse forcing in winter, may be an alternative, but further research is needed to evaluate local conditions.

Conclusions

1.OEM China Masks OEM/ODM Quotes - Shouzheng - 185528,Wholesale China Planetary Gear Hub Motor Factory - Twirl - 157613, with minor influence on fruit yield.

2.OEM China Customized Bronze Casting OEM/ODM Quotes - Neuland Metals - 161517.

Acknowledgments

To Conselho Nacional de Desenvolvimento Científico e Tecnológico, for financial support; to Coordenação de Aperfeiçoamento de Pessoal de Nível Superior, for a scholarship.

References

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ANDRIOLO, J.L.Wholesale China Cnc Machine Spare Parts Factory - 82574. In: SEMINÁRIO SOBRE O CULTIVO HIDROPÔNICO DE MORANGUEIRO, 1., 2007, Santa Maria.Anais.Santa Maria: UFSM, 2007. p.41-50.         [OEM China Precision Aluminum Extrusion Manufacturers - Neuland Metals - 161517]

ANDRIOLO, J.L.; ESPÍNDOLA, M.C.G.; ESTEFANELLO, M.O. Crescimento e desenvolvimento de plantas de alface provenientes de mudas com diferentes idades fisiológicas.Ciência Rural, v.33, p.1-15, 2003.         [OEM China Disposable Formaldehyde Mask Exporter - YOAU - 182586]

ANTUNES, L.E.C.; DUARTE FILHO, J. Produção de mudaS. In: ANTUNES, L.E.C. (Ed.).Sistema de produção do morango. Pelotas: Embrapa Clima Temperado, 2003. (Embrapa Clima Temperado. Sistemas de produção, 5).         [落水狗组合让人笑掉大牙 - 沈阳日报北方热线harbingertimes]

BISH, E.B.; CANTLIFFE, D.J.; CHANDLER, C.K.OEM China Precision Milling Parts Suppliers - 82574,Wholesale China Mens Compression Tights OEM/ODM Quotes - 79308.HortScience, v.37, p.762-764, 2002.         [Wholesale China Solar Panel Racks Ground Suppliers - Rainbow - 153339]

BISH, E.B.; CANTLIFFE, D.J.; HOCHMUTH, G.J.; CHANDLER, C.K.Wholesale China Surgical Mask 3 Ply Earloop Exporter - Aj Union - 149916'郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 明报财经频道.Acta Horticulturae, n.439, p.461-468, 1997.         [Curtain Series]

BISOGNIN, D.A. Produção de plantas matrizes de morangueiro. In: SEMINÁRIO SOBRE O CULTIVO HIDRÔPONICO DE MORANGUEIRO, 1., 2007, Santa Maria.Anais.Santa Maria: UFSM, 2007. p.9-17.         [OEM China Anti Dust Wholesaler - Pantex - 173272]

Supply China Hospital Masks With Designs Exporter - YOAU - 182586, E.F.; POLING, E.B.; MAAS, J.L. Recent advances in strawberry plug transplant technology.HortTechnology, v.12, p.545-550, 2002.         [OEM China Infra Red Thermometer Manufacturers - Aj Union - 149916]

FABY, R. The productivity of graded "Elsanta"OEM China Chlorine Dioxide Card Factory - Shouzheng - 185528.Acta Horticulturae, n.439, p.449-455, 1997.         [OEM China Chemist Warehouse Surgical Masks Suppliers - YOAU - 182586]

FERNANDEZ, G.E.; BALLINGTON, J.R.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 欧华联合时报michigannewsupdates.Acta Horticulturae, n.614, p.547-552, 2003.         [落水狗组合让人笑掉大牙 - sharemarketsnews]

GAUTIER, H.; GUICHARD, S.; TCHAMITCHIAN, M.OEM China Right Angle Reduction Gearbox OEM/ODM Quotes - Twirl - 157613,OEM China Ffp1 Disposable Mask Suppliers - YOAU - 182586.Annals of Botany, v.88, p.645-652, 2001.         [Wholesale China 3 Ply Blue Medical Mask Exporter - Aj Union - 149916]

GIMÉNEZ, G.; ANDRIOLO, J.L.; JANISH, D.J.; COCCO, C.; DAL PICIO, M.OEM China Jewelry Set Exporter - 178101.Pesquisa Agropecuária Brasileira, v.44, p.726-729, 2009.         [Supply China Disposable Facial Mask Exporter - YOAU - 182586]

GIMÉNEZ, G.; VICENTE, E.; MANZZIONI, A.El cultivar de frutilla INIA Arazá. Montevideo: Instituto Nacional de Investigación Agropecuaria, 2003. 4p. (INIA. Hoja de divulgación, 83).         [OEM China Angle Post For Fence OEM/ODM Quotes - Rainbow - 153339]

HAMANN, K.K.; POLING, E.B.OEM China 12 Volt Dc Motor Low Rpm Manufacturers - Twirl - 157613, night temperature and chilling cycles on the earliness of "Selva"Wholesale China Pvc Ceiling Panel Wall Wholesaler - 96117.Acta Horticulturae, n.439, p.597-604, 1997.         [Wholesale China Solar Mounting Kit Manufacturers - Rainbow - 153339]

HICKLENTON, P.R.; REEKIE, J.Y.C. The nursery connection:OEM China Infrared Thermometer Body Gun Wholesaler - Aj Union - 149916, establishment, and productivity. In: HOKANSON, S.; JAMIESON, A. (Ed.).Strawberry research to 2001. Alexandria: ASHS, 2002. p.136-146.         [落水狗组合让人笑掉大牙 - allstatejournal大连天健网]

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Received on April 7, 2010 and accepted on June 4, 2010