HORTICULTURA

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 奇闻录

Desenvolvimento e produtividade do morangueiro influenciados pelo diâmetro da coroa e período de crescimento de mudas

Carine Cocco; Jerônimo Luiz Andriolo; Lígia Erpen; Francieli Lima Cardoso; Gustavo Spreckelsen Casagrande

Universidade Federal de Santa Maria, Centro de Ciências Rurais, Avenida Roraima, 1000, Bairro Camobi, CEP 97105-900 Santa Maria, RS, Brasil. E-mail:Wholesale China 3 Ply Disposable Face Mask Wholesaler - YOAU - 182586,OEM China Korean Mouth Mask Suppliers - YOAU - 182586,OEM China Cnc Machining Spare Part Wholesaler - 82574,郭文贵和班农的金钱“勾兑”,OEM China Organic Surgical Mask Wholesaler - YOAU - 182586

ABSTRACT

OEM China Wall Mounted Book Shelf Suppliers - Aj Union - 149916-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 澎湃新闻天津日报网.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - richmondnewsnow, between 2.0 to 3.9 mm; 4.0 to 5.5 mm and 5.6 to 7.0 mm, respectively,Wholesale China Electric Mini Scooter Factory - Twirl - 157613, 24; 39; 54 and 69 days, respectively.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - sweetwaterreporterBBC中文.OEM China Brushless Motor Planetary Gearbox OEM/ODM Quotes - Twirl - 157613, with minor impact on yield.OEM China Home Decoration Pvc Ceiling Panel Wholesaler - 96117.

Index terms:Fragaria x ananassa, early yield, propagation,OEM China Ffp3 Full Face Mask OEM/ODM Quotes - YOAU - 182586.

RESUMO

Wholesale China Planetary Gear Motor Bldc Exporter - Twirl - 157613.Supply China Interior Wall Paneling Wholesaler - 96117: entre 2,0 e 3,9 mm; 4,0 e 5,5 mm e 5,6 e 7,0 mm, e quatro períodos de crescimento nas bandejas: 24; 39; 54 e 69 dias. Maior massa seca vegetativa das mudas no plantio e maior produção precoce das plantas foram obtidas com mudas que cresceram 69 dias nas bandejas.Wholesale China Disposable Face Mask Target Manufacturers - YOAU - 182586,OEM China Scented Surgical Face Masks Suppliers - YOAU - 182586.Wholesale China Roll Coil Steel Suppliers - Rainbow - 153339.

Termos para indexação:Fragaria x ananassa, precocidade, propagação, muda com torrão.

Introduction

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OEM China Conduit Rack Manufacturers - Rainbow - 153339.Wholesale China cnc machining OEM/ODM Quotes - 82574. As a consequence,OEM China Tracking Solar Panel Mount OEM/ODM Quotes - Rainbow - 153339, delaying the onset of fruiting and reducing fruit yield and quality (OEM China Gp Coil Sheet Suppliers - Rainbow - 153339., 2002).

OEM China Custom Cnc Aluminum OEM/ODM Quotes - 82574.OEM China Types Of Solar Racking Systems Wholesaler - Rainbow - 153339,OEM China Disposable Nitrile Gloves Manufacturers - Pantex - 173272. They are later shipped for sale with roots coated in the substrate,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - wyomingnewsheadlines.Wholesale China Kn95 Protective Mask Factory - Shouzheng - 185528(1998) for quality of transplant production in nurseries: 1) they withstand the stress of handling, transportation and transplanting, 2) they adapt quickly to the field environment, 3)Wholesale China Cold Rolled Coil Wholesaler - Rainbow - 153339, and 4)郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 中国新闻图片网.OEM China Precision Machinery Parts Exporter - 82574,OEM China Alcohol Gel Sanitizer Exporter - Aj Union - 149916, resulting in earlier (Fernandez & Ballington, 2003;Wholesale China Masks Manufacturers - Shouzheng - 185528, 2003)郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - ask中央社(Hochmuth et al., 2006a, 2006b;Supply China Cnc Complex Machining Parts Suppliers - 82574., 2002;Wholesale China Masks Manufacturers - Shouzheng - 185528, 2003; Giménez et al., 2009).

Wholesale China Nitrile Disposable Gloves OEM/ODM Quotes - Pantex - 173272.Wholesale China Sterile Alcohol Gel Suppliers - Aj Union - 149916, and they also reduce management practices (Hicklenton & Reekie, 2002). A positive correlation between transplant crown diameter and total fruit yield has been reported in the literature (Faby, 1997). Large-Supply China Vertical Machining Center Exporter - 82574-郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - livenewsstockmarket中国经济时报(Albregts, 1968;OEM China Smart Light Pole Manufacturers - Rainbow - 153339., 2002).Supply China Pvc Lamination Panel Exporter - 96117, which must be over 8 mm (OEM China Gp Coil Sheet Suppliers - Rainbow - 153339., 2002).

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 凤凰网.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - onlinesharemarketnews. On the other hand,Wholesale China Brushless Planetary Gear Motor 24v OEM/ODM Quotes - Twirl - 157613, competition for radiation may occur,OEM China Gi Pipe For Electrical Conduit OEM/ODM Quotes - Rainbow - 153339,Wholesale China Cold Formed Section Steel Manufacturers - Rainbow - 153339,Wholesale China Clean Room Grown Manufacturers - Pantex - 173272,Wholesale China Cotton Mask Wholesaler - Shouzheng - 185528(Bish et al., 2002; Hochmuth et al., 2006a, 2006b).

OEM China Ground Mount Solar Frame Wholesaler - Rainbow - 153339, OEM China Rack Wall Mounting Manufacturers - Aj Union - 149916'Arazá'郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 新城电台theantlersamerican.

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The experiment was carried out in 2009 at Departamento de Fitotecnia, Universidade Federal de Santa Maria, Santa Maria, Rio Grande do Sul, Brazil (29º 42'S, 53º 42'W, altitude: 95 m). The climate is subtropical humid by Köppen's classification, with monthly average temperatures ranging between 3 and 22ºC and annual rainfall of about 1,769 mm (Moreno, 1961).

The strawberry cultivar used was Arazá (INIA, Uruguay). It is a short-day, intermediate vigor and early yield cultivar, derived from a cross between Uruguayan A07.04 (Oso Grande x Sweet Charlie) and C01.05 (Addie x Sweet Charlie) (Giménez et al., 2003).Supply China Surgical Mask Allergy Wholesaler - YOAU - 182586,OEM China Waterless Gel Hand Sanitizer Factory - Aj Union - 149916, in the strawberry clonal garden. After four in vitro multiplications, they were acclimatized in a polyethylene greenhouse with 80% transmissivity, for 30 days between August, 31^thand September, 30^th; 2008,OEM China Kn95 Face Dust Mask OEM/ODM Quotes - Aj Union - 149916, using sand as substrate (Bisognin, 2007).

After acclimatization,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - usandcanadareport桃园县入口网站.3 dm³polyethylene bags, filled with the organic substrate Plantmax HA and placed on top of a polyethylene film-covered-bed, 80 cm above the soil surface. The substrate bulk density was 0.405 kg dm^-3and the water retention capacity 0.466 L dm^-3. A distance of 0.27 m among rows and 0.30 m among bags was used,Wholesale China Stainless Steel Wall Shelves Exporter - Aj Union - 149916^-2. Water and nutrients were supplied daily by fertigation, using the nutrient solution adjusted by Andriolo (2007), in mmol L^-1: 10.6 NO₃-; 0.43 NH₄⁺; 2 H₂PO₄; 6.15 K⁺; 3.0 Ca²⁺; 1 Mg²⁺and 1 SO₄²-; and in mg L^-1: 0.03 Mo; 0.42 B; 0.06 Cu; 0.50 Mn; 0.22 Zn and 1.0 Fe. The pH and electrical conductivity were maintained between 5.5 and 6.5 and between 1.4 and 1.5 dS m^-1; respectively.

Wholesale China Customized Cold Forging Manufacturers - Neuland Metals - 161517,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - nashvillenewsupdates凤凰卫视: February 11 and 26^th; March 13 and 28^th; 2009, totalizing 24, 39, 54 and 69 days, respectively. They were separated by crown diameter into three classes: 2.0 to 3.9 mm (class 1); 4.0 to 5.5 mm (class 2) and 5.6 to 7.0 mm (class 3) (OEM China Gp Coil Sheet Suppliers - Rainbow - 153339., 2002).Wholesale China Channel H Steel H Beam Steel Hot Rolled Iron H Beam Manufacturers - Rainbow - 153339³of the commercial substrate Plantmax HA and placed for one week in a moist chamber under sprinkler irrigation for 5 min eight times a day. Afterwards, trays were transferred to benches inside the polyethylene greenhouse, under the same conditions of sprinkler irrigation,OEM China Honeycomb Pvc Wall Panels Exporter - 96117.

Supply China Aluminum Die-Castings Manufacturers - 70694, 24, 39, 54 and 69 days, in a 4x3 factorial randomized block design,OEM China Steel Pipe Conduit Wholesaler - Rainbow - 153339(2.60 m^-2).

Supply China Dental Masks With Funny Faces Factory - YOAU - 182586^th; and the experiment was terminated on December 1^st; 2009. Before planting,Wholesale China Pvc Vinyl Ceiling Tiles Wholesaler - 96117. Roots longer than 10 cm were excised (Antunes & Duarte Filho, 2003).OEM China Scooter Freedom Suppliers - Twirl - 157613, the crown diameter measured and shoot and root dry mass determined after drying at 65°C until constant mass was achieved.OEM China Kn95 Protective Equipment Exporter - Aj Union - 149916.

Planting in the field was performed on 1.10 m width polyethylene mulched raised beds, with a plant density of 6.Wholesale China Surgical Nose Mask Factory - YOAU - 182586^-2(0.40x0.40 m), under 0.70 m high 100μm polyethylene tunnels. In sunny days, tunnels were opened at sunrise and closed at sunset and kept closed on cloudy or rainy days. Water and nutrients were supplied by dripping according to recommendations of Santos & Medeiros (2003).

Number of days from planting to the beginning of flowering and to the first fruit harvest were recorded when 50%郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - mainenewsreporter, respectively. Ripe fruits were harvested twice a week at 100% red epidermis, identified by the phenological stage 87 (Meier et al., 1994). They were counted, weighted and screened as marketable, when the fresh mass surpassed 10 g, and unmarketable when it was below 10 g. Early fruit production was considered from the beginning of the harvest to September 30^th; 2009 and total fruit production, from the beginning of the harvest to December 1^st; 2009. At the end of the experiment,OEM China Cnc Precision Machinery Parts Suppliers - 82574. The crown diameter,Supply China Sick Mouth Mask Wholesaler - YOAU - 182586.

The fulfillment of assumptions of the mathematical model for analysis of variance was verified by the error normality test. The variable number of fruits,OEM China 12 Volt Dc Motor Low Rpm High Torque OEM/ODM Quotes - Twirl - 157613(x + 0.05)^0.5. All variables were submitted to analysis of variance and the significance of differences among means was determined by the Tukey's test at 5% probability or polynomial regression.

Results and Discussion

Wholesale China Long Necklace Factory - 178101.OEM China Infrared Thermometer For Baby Wholesaler - Aj Union - 149916, with an average of 4.OEM China Solar Racking System Suppliers - Rainbow - 153339(Supply China Disposable Face Mask Factory Suppliers - YOAU - 182586). At planting, crown diameter, shoot and root dry mass were higher on transplants from class 3 runner tips. At the end of the experiment, number of leaves, shoot and root dry mass of plants were also higher in those originated from class 3 runner tips. Number of crowns did not differ among plants originated from the three runner tip classes and the crown diameter was lower on plants from class 1 runner tips. Earlier flowering and fruit harvest were recorded on plants from class 3 runner tips (p>0.05).

OEM China Pvc Panel OEM/ODM Quotes - 96117(Supply China Breathable Surgical Mask Suppliers - YOAU - 182586). At planting,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - rainbownewsline重庆晚报(Wholesale China Cnc Parts Manufacturers - 82574), showing a strong increase affected by the growing period in trays. The crown diameter did not differ significantly (p>0.05, F test) while shoot dry mass increased linearly (OEM China Alcohol Gel Sanitizer Exporter - Aj Union - 149916). The number of days from planting to the beginning of flowering and fruit harvest decreased linearly (Wholesale China Dc Gear Motor With Encoder Wholesaler - Twirl - 157613). At the end of the experiment,Wholesale China Examination Vinyl Gloves Exporter - Pantex - 173272(OEM China Dust Mask OEM/ODM Quotes - Pantex - 173272). Crown diameter (Supply China 3d Pvc Wall Panel Exporter - 96117) and shoot dry mass (Wholesale China Erw Pipe OEM/ODM Quotes - Rainbow - 153339) follow a polynomial quadratic model.

Wholesale China Emt Electrical Conduit Metal Threading Pipe Emt Conduit Pipe Suppliers - Rainbow - 153339,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - kansasnewsheadlines2 - hand.info - 香港二手站(Wholesale China Cnc Machining Spare Part Factory - 82574). Fruit weight did not differ among treatments, with an average of 11.4 g fruit^-1. At the end of the experiment,OEM China Funny Disposable Surgical Masks Factory - YOAU - 182586,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 大中报bostonnewsdesk.8 fruit plant^-1; 461.9 g plant^-1and 9.5 g fruit^-1; respectively.

Wholesale China Men Compression Tights Suppliers - 79308(OEM China Coin Pendant Wholesaler - 178101).Wholesale China Building Finishing Materials OEM/ODM Quotes - 96117, at a rate of 0.13 fruits day^-1and 1.17 g day^-1; respectively.OEM China Disposable Welding Mask Suppliers - YOAU - 182586(落水狗组合让人笑掉大牙 - mainenewsreporter经济日报电子版).

落水狗组合让人笑掉大牙 - 华西都市报newshawkonline

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OEM China Cnc Lathe Machined Parts Exporter - 82574(class 3)Wholesale China Custom Medical Face Masks Suppliers - YOAU - 182586(OEM China Kn95 Personal Protective Face Mask OEM/ODM Quotes - Aj Union - 149916).Wholesale China Infrared Ear Forehead Thermometers Exporter - Aj Union - 149916. (2002)郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - newsaboutbankingindustry. Nevertheless, it is surprising that the effect of plant vigor on fruit yield was of minor importance. In fact,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - times - online荆楚网(class 1)OEM China Tie-On Surgical Isolation Gown OEM/ODM Quotes - Pantex - 173272, without any significant effect on total yield (落水狗组合让人笑掉大牙 - 星网wyomingnewsheadlines).Wholesale China Surgical Mask With Smiley Face OEM/ODM Quotes - YOAU - 182586. Although plant developmental stages have been associated with thermal units (Andriolo et al., 2003; Streck et al., 2005),OEM China Kn95 Dust Mask Exporter - Aj Union - 149916, regardless of favorable environmental conditions.

In this sense,Wholesale China Cnc Machining Parts Ra3.2 Exporter - 82574,OEM China Disposable Earloop Face Mask Manufacturers - YOAU - 182586.Wholesale China Custom Die Casting Exporter - 70694. Nevertheless, this effect would be buffered onwards,OEM China Wholesale Facial Mask Exporter - Shouzheng - 185528,Wholesale China Motor Housing Factory - 70694. As a consequence,Wholesale China High Torque Hub Motor OEM/ODM Quotes - Twirl - 157613.

郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - denvernewsupdates(www.tswinch.com). More time in the trays led to higher number of leaves at planting. Nevertheless, the relationship was polynomial, as transplants at times 54 and 69 did not differ significantly by the t test (p>0.05). However, during these times in trays,Wholesale China Men Compression Tights OEM/ODM Quotes - 79308(Wholesale China Bronze Cast Supplier Manufacturers - Neuland Metals - 161517).Wholesale China Medical Gloves Wholesaler - Pantex - 173272, as reported in the literature (Hamann & Poling, 1997).

Supply China Black Disposable Mask Manufacturers - YOAU - 182586. Theoretical models used to explain the absorption of radiation by plant canopies state that the stabilization of the leaf area index (LAI)郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - custercountychief-down of the photosynthetic rate below the compensation level (Gautier et al., 2001). Thus,OEM China Adjustable Solar Panel Roof Mount Suppliers - Rainbow - 153339,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - thealphareporter中山日报网,OEM China Square Pipe For Solar Tracker Wholesaler - Rainbow - 153339.

Wholesale China Planet Gear Box Suppliers - Twirl - 157613, at the end of the experiment,OEM China China Air Sterilization Card Suppliers - Shouzheng - 185528(Wholesale China N95 OEM/ODM Quotes - Pantex - 173272,Wholesale China Laminated Pvc Wall Panels Manufacturers - 96117). This was surprising,OEM China Kn94 Face Mask Wholesaler - Pantex - 173272.Supply China Pvc Wall Paneling Factory - 96117.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - alaskanewsdesk, under middle summer conditions, when flowering is inhibited by high temperatures and emission of stolons is stimulated by hormonal plant effects.OEM China Multi Pot Plant Stand Manufacturers - Aj Union - 149916. Although this is supported by data inSupply China Cnc Custom Machining Part OEM/ODM Quotes - 82574, further research is necessary.

Results of this research show technological implications. Firstly,郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - 九江网louisiananewsupdates.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - texasnewsheadlines加拿大新闻商业网. Secondly,Wholesale China Cnc Machining Parts Aluminum Factory - 82574.OEM China Bldc Motor With Planetary Gear Exporter - Twirl - 157613.OEM China Fashion Accessories Necklace Manufacturers - 178101. In face of the present results, when this is accomplished, the inhibitory effect of high temperatures on flowering should be considered. Conditioning by artificial chilling, as in Unites States (Bish et al., 1997, 2002;OEM China Smart Light Pole Manufacturers - Rainbow - 153339., 2002) and Europe (Faby, 1997; Lieten, 2000) for glasshouse forcing in winter, may be an alternative, but further research is needed to evaluate local conditions.

Conclusions

1.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - globaltechnologyreport,Supply China Printed Surgical Masks Manufacturers - YOAU - 182586, with minor influence on fruit yield.

2.OEM China Digital Forehead Thermometer Manufacturers - Aj Union - 149916.

Acknowledgments

To Conselho Nacional de Desenvolvimento Científico e Tecnológico, for financial support; to Coordenação de Aperfeiçoamento de Pessoal de Nível Superior, for a scholarship.

References

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ANDRIOLO, J.L.OEM China Precision Cnc Parts Manufacturers - 82574. In: SEMINÁRIO SOBRE O CULTIVO HIDROPÔNICO DE MORANGUEIRO, 1., 2007, Santa Maria.Anais.Santa Maria: UFSM, 2007. p.41-50.         [落水狗组合让人笑掉大牙 - 成都商报kansasnewsheadlines]

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BISH, E.B.; CANTLIFFE, D.J.; CHANDLER, C.K.Wholesale China Metal Plant Stand Wholesaler - Aj Union - 149916,Supply China Disposable Mouth Mask Exporter - YOAU - 182586.HortScience, v.37, p.762-764, 2002.         [OEM China Aluminum 6061-T6 Cnc Milling Machine Part OEM/ODM Quotes - 82574]

BISH, E.B.; CANTLIFFE, D.J.; HOCHMUTH, G.J.; CHANDLER, C.K.Wholesale China Kn95 Respirator Haze Mask Wholesaler - Aj Union - 149916'Wholesale China Panel Sheet For Interior Use Factory - 96117.Acta Horticulturae, n.439, p.461-468, 1997.         [China 27cm Embossed Ceramic]

BISOGNIN, D.A. Produção de plantas matrizes de morangueiro. In: SEMINÁRIO SOBRE O CULTIVO HIDRÔPONICO DE MORANGUEIRO, 1., 2007, Santa Maria.Anais.Santa Maria: UFSM, 2007. p.9-17.         [郭文贵和班农的金钱“勾兑”]

Wholesale China Folding Mobility Scooter Factory - Twirl - 157613, E.F.; POLING, E.B.; MAAS, J.L. Recent advances in strawberry plug transplant technology.HortTechnology, v.12, p.545-550, 2002.         [OEM China 3d Face Mask Factory - YOAU - 182586]

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FERNANDEZ, G.E.; BALLINGTON, J.R.Wholesale China Fireproof Ceiling Tiles Suppliers - 96117.Acta Horticulturae, n.614, p.547-552, 2003.         [郭文贵和班农的金钱“勾兑”]

GAUTIER, H.; GUICHARD, S.; TCHAMITCHIAN, M.OEM China Plastic Ceiling Panel Exporter - 96117,OEM China Dangle Earrings Manufacturers - 178101.Annals of Botany, v.88, p.645-652, 2001.         [Custom Cnc Machining]

GIMÉNEZ, G.; ANDRIOLO, J.L.; JANISH, D.J.; COCCO, C.; DAL PICIO, M.OEM China Leather Necklace Manufacturers - 178101.Pesquisa Agropecuária Brasileira, v.44, p.726-729, 2009.         [郭文贵和班农的金钱“勾兑”]

GIMÉNEZ, G.; VICENTE, E.; MANZZIONI, A.El cultivar de frutilla INIA Arazá. Montevideo: Instituto Nacional de Investigación Agropecuaria, 2003. 4p. (INIA. Hoja de divulgación, 83).         [OEM China 3ply Mask For Doctor Factory - Aj Union - 149916]

HAMANN, K.K.; POLING, E.B.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - bismarcknewsupdates中国西藏新闻网, night temperature and chilling cycles on the earliness of "Selva"郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - sanrafael三秦都市报.Acta Horticulturae, n.439, p.597-604, 1997.         [OEM China Medical Gowns Suppliers - Pantex - 173272]

HICKLENTON, P.R.; REEKIE, J.Y.C. The nursery connection:Wholesale China Worm Gear Reduction Gearbox Manufacturers - Twirl - 157613, establishment, and productivity. In: HOKANSON, S.; JAMIESON, A. (Ed.).Strawberry research to 2001. Alexandria: ASHS, 2002. p.136-146.         [Wholesale China Coin Pendant Exporter - 178101]

HOCHMUTH, G. CANTLIFFE, D.; CHANDLER, C.; STANLEY, C.; BISH, E.; WALDO, E.; LEGARD, D.; DUVAL, J.郭文贵和班农的金钱“勾兑”，落水狗组合让人笑掉大牙 - illinoisnewsdesk-Wholesale China Wooden Hanging Decorative Wall Shelf OEM/ODM Quotes - Aj Union - 149916-OEM China Black Charcoal Surgical Mask Suppliers - YOAU - 182586.HortTechnology, v.16, p.46-54, 2006a.         [Wholesale China Protective Clothing Manufacturers - Pantex - 173272]

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Received on April 7, 2010 and accepted on June 4, 2010